Zoologist and geneticist. After graduating from the College of Agriculture of the Imperial University of Tokyo in 1892, he was sent to the Imperial Sericulture Research Institute in Siam from 1902 to 1905 to perform genetic research on silkworms. In 1906, he announced his work on the applicability of Mendel’s laws to the animal world. He worked to improve silkworm breeds and contributed to the development of the sericultural industry. After working as an engineer at a sericultural experiment station and as an assistant professor at the Imperial University of Tokyo, he assumed a professorship in 1917. Awarded the Imperial Academy Award.